Lesaro was a lieutenant in the Spanish Royal Navy during the Age of Piracy, serving on the Silent Mary under the command of Capitán Armando Salazar. Lesaro was the most loyal officer in Salazar's crew, following his captain in his numerous pirate-hunting expeditions. Lesaro's loyalty eventually cost him his life when their ship sailed into the Devil's Triangle where the entire crew was killed in an explosion. However, Lesaro was resurrected by the supernatural powers of the Triangle with his crewmates as the undead. Hungry for revenge, the Spanish escaped from the Triangle and set out on a blind mission of vengeance to find the legendary Trident of Poseidon to kill the man they blamed for their fate - Captain Jack Sparrow. He helped his captain in his quest and regained his human form following the Trident's destruction, only to be devoured by the sea and drown.